Evelyn Fox Keller, a theoretical physicist, a mathematical biologist and, beginning in the late 1970s, a feminist theorist who explored the way gender pervades and distorts scientific inquiry, died on Sept. 22 at an assisted living home in Cambridge, MA. She was 87,

Like many women in the sciences, she had faced years of disparagement and discrimination, and one of her first efforts was to quantify the effect such a hostile environment had on women — how it held them back, and how it drove many to leave science completely.

She pressed her argument in books, essays and speeches, often to packed auditoriums.